The prerequisites for the different chapters change considerably between the chapters themselves. As the third chapter took quite a while to get unlocked, we are able to hop into the fourth immediately after escaping the affinity mission.
For this one, a lifehold has been identified in Noctilim that we have to head out to investigate. Here we manage to find yet another alien species and one of them ends up joining the group. Said individual is the best tank in the game, so there are a lot of strategies you're able to use with him. The problem is that he uses the same shield weapons as Lin, and with her being required on most missions, it becomes difficult to squeeze them both in.