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If \( \mathrm{N} \) is the population density at time \( \mathrm{t} \), then its
P density at time \( t+1 \) is

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(1) \( \mathrm{N}_{\mathrm{t}+1}=\mathrm{N}_{\mathrm{t}}+[(\mathrm{B}+\mathrm{I})+(\mathrm{D}+\mathrm{E})] \)
(2) \( \mathrm{N}_{\mathrm{t}+1}=\mathrm{N}_{\mathrm{t}}-[(\mathrm{B}+\mathrm{I})+(\mathrm{D}+\mathrm{E})] \)
(3) \( \mathrm{N}_{\mathrm{t}+1}=\mathrm{N}_{\mathrm{t}}+[(\mathrm{B}+\mathrm{I})-(\mathrm{D}+\mathrm{E})] \)
(4) \( \mathrm{N}_{\mathrm{t}+1}=\mathrm{N}_{\mathrm{t}}-[(\mathrm{B}+\mathrm{I})-(\mathrm{D}+\mathrm{E})] \)
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