Hyrule Warriors Definitive Edition Walkthrough Part 62: Hard Mode Skulltulas Part 4
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Land of Twilight: Play as Lana using the Book of Sorcery. Defeat Midna before she reaches Cia and capture all keeps to make the Skulltula appear.
The Shadow King: Play as Midna and defeat the three Keep Saboteurs without Lana and Agitha's health going below 50% to make the Skulltula appear.
Land in the Sky: Play as Link using the Hylian Sword and activate the Fairy Circle without Fi's health going below 50% to make the Skulltula appear.
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