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In Warhammer 40,000: Space Marine II, the player engages enemies using a mix of both melee and ranged combat from a third-person point of view. The player controls Titus, a lieutenant of the Ultramarines chapter and returning protagonist from the first game. Titus is accompanied by fellow Space Marines Chairon and Gadriel throughout the campaign mode.

Titus is protected by armor. When his armor breaks, his health will drop permanently when he is attacked. Health will only replenish when the player recovers medkits in the game's battlefield. His armor, however, will regenerate after Titus performs an execution attack on an enemy. If a character's health is depleted, the character will drop to the ground and bleed out for 30 seconds, during which other characters can revive them before they die.

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